Inclusive Fitness Formula
Hamilton s inclusive fitness maintains heritable altruism polymorphism through rb c changcao wanga and xin lua b 1 adepartment of ecology college of life sciences wuhan university wuhan 430072 china.
Inclusive fitness formula. Your inclusive fitness is. Louis mo and approved december 12. Altruistic behaviors have costs which are reductions in direct fitness but if they benfit relatives the indirect fitness benefits to the altruist may offset the direct fitness costs. In either case it describes individual reproductive success and is equal to the average contribution to the gene pool of the next generation that.
Hamilton in 1963 and 1964. Fitness often denoted or ω in population genetics models is the quantitative representation of natural and sexual selection within evolutionary biology it can be defined either with respect to a genotype or to a phenotype in a given environment. 27 the derivations were demonstrated to be unrelated by corresponding parts of the two identical formulae for r displaystyle r being derived from the genotypes of. In evolutionary biology inclusive fitness is one of two metrics of evolutionary success as defined by w.
Bill hamilton s 1963 and 1964 inclusive fitness articles are easily the most cited articles in the entire field of behavioral evolution and his work on altruism and kinship spurred endless dissertation projects and hundreds of published articles both theoretical and empirical. Orlove argued that the existence of two unrelated derivations of the formula for reduces or eliminates the ad hoc nature of the formula and of inclusive fitness theory as well. Hamilton s rule hamilton s rule. Haldane in the causes of evolution the theory was later named and developed by british evolutionary biologist william donald hamilton who used inclusive fitness to explain direct reproductive and indirect aided by a relative or a colony member inheritance of genetic traits associated with altruism.
In evolutionary biology inclusive fitness theory is a model for the evolution of social behaviors traits first set forward by w. Strassmann washington university in st. If you want to reach the 80 of people who aren t exercising at a gym then you need to join health clubs around the world that are working to make facilities more accessible to people of all abilities. Personal fitness is the number of offspring that an individual begets regardless of who rescues rears supports them.
Wellness community programming. And bthe institute for advanced studies wuhan university wuhan 430072 china edited by joan e. 5 15 5 8 125 4 25 2 25 10 2 5 12 5 altruism and inclusive fitness. Instead of a trait s frequency increase being thought of only via its average effects on an organism s direct reproduction hamilton argued that its average effects on indirect reproduction via identical copies of the trait in other.
An open letter to the fitness industry on inclusivity.